I came across this little gem just the other morning when deciding to edit a few of the email signatures that I use.
Turns out that Lion moves the default mail location so somewhere different. If you want to edit your mail signatures navigate to..
/Users/YOURUSERNAME/Library/Mail/V2/MailData
There’s plenty of guides out there on how to edit your mail signature in Lion, sadly I no longer have a Lion installation to be able to help but if you have any hints or tips please email in.
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